Figure above :  Wing of  Clythia  picta MEIG (Clythiidae).
The wing-venation of this form is close to the general venational groundplan of cyclorraphic flies.
The Radial Sector is, as in all cyclorraphic flies, two-branched, that is, it consists of two long more or less parallel branches. The cell lying directly below the second branch of the Radial Sector (R5) is called the "fifth radial cell". Posteriorly it is bounded by the anterior main branch of the Media. The latter branch is connected with the second branch of the Radial Sector by the cross-vein  r-m  ( = ta). The position of this cross-vein is here rather specialized because it has shifted somewhat toward the wing-base. The two main branches of the media enclode the  discoidal cell  ( = intermedial cell). Distally this cell is closed by the  intermedial cross-vein  ( = tp).
The posterior medial branch is connected with the anterior cubital branch (CuA) by the cross-vein  m-cu  ( = tb). CuA bends toward the first anal vein (1A) and coalesces with it. The common stalk (CuA + 1A) of these two veins becomes longer in many other species (of Cyclorrapha and others) implying that the enclosed cell is contracting toward the wing-base. This enclosed cell we earlier had -- following HENNIG -- called the "anal cell", but it is better -- following ROHDENDORF -- to call it the  "cubital cell".

In diagram 1 (further down) we indicate a number of 'character movements' that can formally be distinguished between the venational patterens of some cyclorraphic flies (and a few orthorraphic ones). In fact we observe either a constancy of the state of a given venational character, or we see a change of the state (i.e. a transformation from a plesiomorphous state to an apomorphous state) of a given venational character. For example, the (plesiomorphous) state of the cubital cell has changed (into a more apomorphous state) when the cell's contraction (toward the wing-base) has proceeded further. The changed -- apomorphous -- state is indicated, in the mentioned diagram, by a blue point at the end of the (red) line connecting this state with the initial -- plesiomorphous -- state (of the same character). A red line without such a blue point connects the same states of the given character :

In all this we assume that there cannot take place transitions that go from an apomorphic to a plesiomorphic state of a given character, that is, evolution will never be (exactly) reversed.

In the diagram that follows, that is, in the mentioned Diagram 1, wings of a number of (recent) cyclorraphic (and some other) species of flies are compared with each other as to their wing-venation :

A -- Nephrocerus  scutellatus MACQ.  ( Dorilaidae [= Pipunculidae] ) (Cyclorrapha). (After HENNIG 1954),
B -- Syrphus  ribesii L.  (Syrphidae) (Cyclorrapha). (After HENNIG 1954),
C -- Conops  flavipes L.  (Conopidae) (Cyclorrapha). (After HENNIG 1954),
D -- Sicus  ferrugineus L.  (Conopidae) (Cyclorrapha). (After HENNIG 1954)
E -- Clythia  picta MEIG.  (Clythiidae) (Cyclorrapha). (After ROHDENDORF 1951),
F -- Callimyia  speciosa MEIG.  (Clythiidae) (Cyclorrapha). (After HENNIG 1954),
G -- Sciopus  albifrons MEIG.  (Dolichopodidae) (Orthorrapha). (After HENNIG 1954),
H -- Dolichopus  picipes MEIG.  (Dolichopodidae) (Orthorrapha). (After HENNIG 1954),
I -- Fannia  sp.  (Anthomyiidae) (Cyclorrapha). (After ROHDENDORF 1951),
J -- Orthellia  sp.  (Muscidae) (Cyclorrapha). (After ROHDENDORF 1951),
K -- Linnaemyia  vulpina FALL.  (Larvaevoridae) (Cyclorrapha). (After ROHDENDORF 1951).

Comparison of the states of certain venational characters -- as present in the venational groundplan of the Cyclorrapha, see figure above -- is expressed in terms of  noëtic interactions  (taking place in the Implicate Order, and the results projected into the Explicate Order).

Diagram 1 :  Morphological relations between the states of a number of characters as they are found in the wing-venation of recent cyclorraphic (and a few other) flies :

• The (plesiomorphous) state of the cubital cell in  A  (Nephrocerus, Dorilaidae) is also found in  B  (Syrphus, Syrphidae). This might be the result of noëtic interaction between the formal contents (as they are, in the form of noëtic entities, present in the Implicate Order) of  Nephrocerus  scutellatus  and  Syrphus  ribesii.
  An evolutionary derivation of the whole wing of  A  from that of  B  (or the other way around) seems unlikely. Instead, only one character has been 'captured' by one of them from the other :  formation of the cubital cell (from a condition in which it did not yet exist).
  
• If we go from  A  (Nephrocerus, Dorilaidae) to  C  (Conops, Conopidae) we see a transformation of the fifth radial cell (loss of M2 and debouching of M1 in posterior branch of Rs). This transformed cell might also have its previous, that is, its plesiomorphous, state in  D  (Sicus, also Conopidae), or even in  B  (Syrphus, Syrphidae).
  
• In  E  (Clythia, Clythiidae) we see the cubital cell still in its primitive state, while the position of  r-m  is apomorphous :  the cross-vein is shifted toward the wing-base. It is apomorph with respect to, for instance,  I  (Fannia, Anthomyiidae).  In  I  (Fannia) the state of the cubital cell is apomophous with respect to that in  E  and  F  (both Clythiidae). But the wing-venation of  I  (Fannia) cannot phylogenetically be derived from that of either  E  or  F  (both Clythiidae) because that would imply a reverse evolutionary course of the state of the cross-vein  r-m ,  that is its position. In the same way  E  or  F  (both Clythiidae) cannot have evolved from  I  (Fannia), or from any other higher cyclorraphous fly for that matter, because that would imply a reverse evolutionary course of the state of the cubital cell.  The straight anterior main branch of the Media in  I  (Fannia) results, not from  I  having been descended from  F  (forbidden by the position of  r-m), but results from having the precursor of  I's formal content in the Implicate Order, or at least that substructure of this (precursor of the) formal content of Fannia that represents its wing-venation, noëtically interacted with the corresponding -- also residing in the Implicate Order -- noëtic substructure in  F  (this substructure representing the fact of the anterior main branch of the Media being simple and more or less straight). So  I  has not descended from  F.  Instead,  I's  precursor has taken up one character only, namely the more or less straight course of the anterior branch of the Media.
  The preservation of the primitive position of the cross-vein  r-m  in  I  (Fannia, Anthomyiidae) could be due to noëtic contact with, say,  J  (Orthellia, Muscidae),  or with  D  (Sicus, Conopidae).

So in addition to the one given venation being, as a whole, evolutionarily derived from some other (whole) venation (which derivation was discussed earlier), some  p a r t s,  and only these parts, of a given venation are derived from the corresponding parts of another venation. And the latter phenomenon is the result of noëtic contact or interaction of the formal contents of two or more (noëtically more or less closely related ) species as these contents are present in the Implicate Order as noëtic entities, structures or patterns.